Great-Grandma-473-08 is a female dognose.
One of the oldest women in the Beagle Boys clan, Great-Gradnma-473-08 spends her old age babysitting the younger generations, such as “grandkids number 378-420 through 378-371, inclusive”. In 1975, when news broke out that the Duckburg trio of Beagles had briefly acquired legal ownership of Scrooge McDuck's fortune, Great-Grandma and the grandkids were among the hundreds of Beagle relatives who suddenly dropped in to mooch off of them.
Behind the scenes
Great-Grandma-473-08 appears in the 1975 story Scrooge For A Day.